
Green (algal) waste from Galápagos marine iguana (Amblyrhynchus cristatus). Isla Fernandina, Galápagos.

Galápagos marine iguanas (Amblyrhynchus cristatus) basking on lava. Isla Fernandina, Galápagos.

Mess of marine iguanas (Amblyrhynchus cristatus) on beach. Isla Fernandina, Galápagos.

Galápagos marine iguana (Amblyrhynchus cristatus) with Sally Lightfoot crabs. Isla Fernandina, Galápagos.

Dead marine iguana (Amblyrhynchus cristatus) being eaten by Sally Lightfoot crabs. Isla Fernandina, Galápagos.

Marine iguana (Amblyrhynchus cristatus) in field of Galápagos carpetweed (Sesuvium edmonstonei). Cerro Dragon, Isla Santa Cruz, Galápagos.

Marine iguana (Amblyrhynchus cristatus) sunning on guano-covered rock. Galápagos Islands, Ecuador.

Marine iguana (Amblyrhynchus cristatus) resting in tide pool. Isla Santiago (James Island), Galápagos, Ecuador.

Juvenile Galápagos marine iguana (Amblyrhynchus cristatus). Isla Fernandina, Galápagos.

Pink marine iguana (Amblyrhynchus cristatus). The marine iguanas on Isla Santiago (formerly James Island) are typically greenish so this individual stood out. Galápagos, Ecuador.

Marine iguana (Amblyrhynchus cristatus) resting in tide pool. Isla Santiago, Galápagos, Ecuador.

Skeletonized marine iguana (Amblyrhynchus cristatus). Isla Santiago, Galápagos, Ecuador.

Galapagos marine iguana (Amblyrhynchus cristatus) sneezing. Punta Espinosa, Fernandina Island, Galapagos.

Fernandina marine iguana (Amblyrhynchus cristatus cristatus) swimming. Isla Fernandina, Galápagos.

Tricuspate teeth on skeleton of a marine iguana (Amblyrhynchus cristatus). Isla Fernandina, Galápagos.

Marine iguana (Amblyrhynchus cristatus) resting in tide pool. Isla Santiago, Galápagos, Ecuador.

Marine iguana (Amblyrhynchus cristatus) on sandy beach. Isla Santiago, Galápagos, Ecuador.

Mess of marine iguanas (Amblyrhynchus cristatus ssp. wikelskii). Puerto Egas, Isla Santiago, Galápagos.

Fernandina marine iguana (Amblyrhynchus cristatus cristatus) yawning. Punta Espinosa, Fernandina Island, Galápagos Island.

School of red cardinal fish (Apogon sp.). Off Isla Genovesa, Galápagos.

Great blue heron (Ardea herodias) on beach. Isla Santa Cruz, Galápagos Islands, Ecuador.

Great blue heron (Ardea herodias) overlooking ocean. Isla Santa Cruz, Galápagos Islands, Ecuador.

Black mangrove (Avicennia germinans) growing in lagoon. Isla Santa Cruz, Galápagos.

Galápagos lava cactus (Brachycereus nesioticus). Isla Fernandina, Galápagos.

Galápagos hawk (Buteo galapagoensis). Isla Fernandina, Galápagos.

Sanderling (Calidris alba). Cero Dragón, Galápagos Islands, Ecuador.

Red drupes on Galápagos bitterbush (Castela galapageia). Isla Isabela, Galápagos.

Baby Galápagos tortoises (Chelonoidis sp.) eating arroleaf elephant’s ear (Xanthosoma sagittifolium) at the Fausto Llerena Tortoise Center in Puerto Ayora, Isla Santa Cruz, Galápagos.

Baby Galápagos tortoises (Chelonoidis sp.) eating arroleaf elephant’s ear (Xanthosoma sagittifolium) at the Fausto Llerena Tortoise Center in Puerto Ayora, Isla Santa Cruz, Galápagos.

Close-up of a western Santa Cruz giant tortoise (Chelonoidis porteri). El Chato Tortoise Reserve near Los Gemelos, Isla Santa Cruz, Galápagos.

Western Santa Cruz Island giant toroise (Chelonoidis porteri). El Chato Tortoise Reserve near Los Gemelos, Isla Santa Cruz, Galápagos.

Western Santa Cruz Island giant tortoise (Chelonoidis porteri). One of two species on the Island. El Chato Giant Tortoise Reserve near Los Gemelos, Santa Cruz, Galápagos.

Juvenile Alcedo Volcano giant tortoise (Chelonoidis vandenburghi). Urbina Bay, Volcan Alcedo. Isla Isabela. Galápagos, Ecuador.

Disarticulating carapace and plastron of a dead Alcedo Volcano giant tortoise (Chelonoidis vandenburghi). Urbina Bay, outskirts of Volcan Alcedo.

Alcedo giant tortoise (Chelonoidis vandenburghi). Urbina Bay, Isla Isabela, Galápagos.

Land iguana (Conolophus subcristatus). Cerro Dragón, Isla Santa Cruz, Galápagos.

Galápagos land iguana (Conolophus subcristatus) eating. Isla Santa Cruz, Galápagos.

Galápagos land iguana (Conolophus subcristatus) near Waltheria ovata. Isla Isabela, Galápagos.

Galapagos land iguana (Conolophus subcristatus). Cerro dragón, Isla Santa Cruz, Galápagos.

Land iguana (Conolophus subcristatus). Isla Isabela, Galápagos.

Pair of courting swallow-tailed gull (Creagrus furcatus). Genovesa, Galápagos.

Pair of swallow-tailed gulls (Creagrus furcatus) under Opuntia. Isla Genovesa, Galápagos.

Dodder (Cuscuta sp.) on goat’s foot Convolulus (Ipomoea pes-caprae). Isla Santiago, Galapagos.

Euphorbia amplexicaulis. Isla Genovesa, Galápagos.

Flock of great frigatebirds (Fregata minor). Genovesa, Galápagos Islands.

Great frigatebird (Fregata minor) showing green tinge on wings. Isla Genovesa, Galápagos.

Great frigatebird (Fregata minor) with inflated gular sac. Genovesa, Galápagos Islands.

Great frigatebird (Fregata minor) perched in shrub. Isla Genovesa, Galápagos.

Great frigatebird (Fregata minor) with inflated gular sac. Genovesa, Galápagos Islands.

Great frigatebird (Fregata minor) in tree. Isla Genovesa, Galápagos.

Galapaganus conwayensis. ID tentative. Isla Santa Cruz, Galápagos.

Evolution’s Workshop, Galápagos: World’s End, Flowering Plants of the Galápagos.

Loathed by local merchants because products are tad more authentic and tourists like them better. Puerto Ayora, Isla Santa Cruz, Galápagos.

Galápagos Islands origin mural at The Rock, Puerto Ayora, Santa Cruz Island, Galápagos.

Galápagos Islands, Ecuador.

Galápagos, Ecuador.

Luxury yacht, 192′, 32 passenger. Anchored off of Isla Santa Cruz, Galápagos.

Puerto Ayora, Isla Santa Cruz, Galápagos.

Galápagos, Ecuado

Galápagos Islands, Ecuador.

Tagus Cove, Isabela Island, Galápagos.

Galápagos, Ecuador.

293′. Galápagos, Ecuador.

Galápagos Islands, Ecuador.

Eroded tuff cone. Isla Bartolomé, Galápagos Islands, Ecuador.

Isla Rabida, Galápagos.

Galápagos Islands, Ecuardor.

Isla Santiago (James Island), Galápagos, Ecuador.

Galápagos, Ecuador.

Isla Santiago (James Island), Galápagos, Ecuador.

Isla Genovesa, Galápagos.

The middle t-shirt has a quote from Leon Megginson, a business professor from Louisiana. But often attributed to Darwin by publishers of motivation posters, t-shirt vendors.

Galápagos Islands, Ecuador.

Common cactus finch (Geospiza scandens) on Opuntia flower. Puerto Ayora, Isla Santa Cruz, Galápagos.

Common cactus finch (Geospiza scandens intermedia) foraging on giant prickly pear (Opuntia echios var. echios) on Isla Santa Cruz, Galápagos, Ecuador.

Gossypium darwinii. Isla Isabela, Galápagos.

Sally Lightfoot crab (Grapsus grapsus) eating another Sally Lightfoot crab. Fernandina Island, Galápagos Islands.

Sally Lightfoot crab (Grapsus grapsus). Fernandina Island, Galápagos Islands.

Sally Lightfoot crab (Grapsus grapsus). Isla Santa Cruz, Galápagos.

Sally Lightfoot crab (Grapsus grapsus). Fernandina Island, Galápagos Islands.

Manchineel (Hippomane mancinella). Isla Isabela, Galápagos.

Galápagos humpback scorpionfish (Idiastion hageyi) on beach. Isla Rabida, Galápagos.

Brown sea cucumber (Isostichopus fuscus) with red cardinal fish. Off Isla Genovesa, Galápagos.

Rustic sphinx moth (Manduca rustica). Isla Santa Cruz, Galápagos.

Fruit of Galápagos leatherleaf (Maytenus octogona). Celastraceae. Isla Santa Cruz, Galápagos.

Galapagos lava lizard (Microlophus albemarlensis) protecting toes from heat. Isla Rabida, Galápagos.

Galápagos lava lizard (Microlophus albemarlensis) on lava. Isla Isabela, Galápagos.

Santiago lava lizard (Microlophus jacobii). Isla Santiago, Galápagos, Ecuador.

Galápagos mockingbird perched on cactus. Puerto Ayora, Santa Cruz Island.

Fledgling Galápagos mockingbird (Mimus parvulus parvulus) in a tree. Charles Darwin Research Station, Puerto Ayora, Isla Santa Cruz, Galápagos, Ecuador.

Black-eared Galápagos mockingbird (Mimus parvulus personatus). Isla Santiago, Galápagos.

Black-eared Galápagos mockingbird (Mimus parvulus personatus) in a tree. Isla Santiago, Galápagos.

Galápagos flycatcher (Myiarchus magnirostris). Isla Santa Cruz, Galápagos Islands, Ecuador.

Galápagos flycatcher (Myiarchus magnirostris) on weathered lava. Isla Santiago (James Island), Galápagos, Ecuador.

Western spotted orbweaver (Neoscona oaxacensis). In the Galápagos this spider is often called (incorrectly) Neoscona cooksoni. The insect caught in the web is probably a yellow paper wasp (Polistes versicolor). Isla Santiago, Galapagos.

Yellow-crowned night heron (Nyctanassa violacea pauper). Isla Genovesa, Galápagos.

Galapagos hawk (Buteo galapagoensis). Isla Genovesa, Galapagos.

Close-up of prickly pear (Opuntia echios gigantea) stem showing spines (modified leaves) and glochids (tufts of short bristles, which are probably modified trichomes). Cerro Dragon, Isla Santa Cruz, Galápagos.

Prickly pear (Opuntia echios gigantea). Cerro Dragón, Isla Santa Cruz, Galápagos.

Fruit of stinking passionflower (Passiflora foetida). Isla Santa Cruz, Galápagos.

Stinking passionflower (Passiflora foetida). Isla Santa Cruz, Galápagos Islands.

A brown pelican (Pelecanus occidentalis urinator) making a wet landing near Fernandina Island, Galapagos.

Brown pelican (Pelecanus occidentalis) with Daphne Minor in background. Black Turtle Cove, Isla Santa Cruz, Galápagos, Ecuador.

Brown pelicans (Pelecanus occidentalis urinator) in flight over ocean. Galápagos, Ecuador.

Brown pelican (Pelecanus occidentalis urinator) landing on water. Near Cabo Nepean, Isla Santiago in the Galápagos.

Brown pelican (Pelecanus occidentalis) in flight. Galápagos, Ecuador.

Ventral view of a brown pelican (Pelecanus occidentalis urinator). Near Cabo Nepean, Isla Santiago, Galápagos, Ecuador.

Brown pelican (Pelecanus occidentalis urinator) landing on water. Near Cabo Nepean, Isla Santiago in the Galápagos.

Brown pelican (Pelecanus occidentalis urinator). Isla Santa Cruz, Galápagos.

Panamic cushion star (Pentaceraster cumingi). Great Darwin Bay, San Cristobal, Galapagos.

Panamic cushion stars (Pentaceraster cumingi). Great Darwin Bay, Isla Genovesa, Galápagos.

Galápagos cormorant (Phalacrocorax harrisi) drying its wing feathers. Fernandina Island, Galápagos Islands.

Galápagos cormorant (Phalacrocorax harrisi) in nest. Fernandina Island, Galápagos Islands.

Blue seastar (Phataria unifascialis). Great Darwin Bay, Isla Genovesa, Galápagos.

American flamingo (Phenicopterus ruber glyphorhynchus). Cero Dragón, Isla Santa Cruz, Galápagos.

American flamingo (Phenicopterus ruber glyphorhynchus) in the Galápagos. Cero Dragón, Isla Santa Cruz, Galápagos.

Variegated paper wasp (Polistes versicolor) nest. Urbina Bay, Isla Isabela, Galápagos.

Elephant bush (Portulacaria afra). At the time, unreported on the Galapagos. I reported it. Isla Rabida, Galápagos.

Skull of a black rat (Rattus rattus). Santiago Island in the Galápagos.

Close-up of coral (Scleractinia) on beach. Darwin Bay Genovesa Island, Galápagos.

Thorn shrub (Scutia spicata var. pauciflora) with fruit. Urbina Bay, outskirts of Volcan Alcedo. Isla Isabela. Galápagos, Ecuador.

Sea purslane (Sesuvium portulacastrum). Cerro Dragón, Isla Santa Cruz, Galápagos.

Yellow warbler (Setophaga petechia aureola) in a tree. Female. Highlands of Santa Cruz Island, Galápagos Archipelago. Sometimes called a mangrove warbler, and formerly classified as Dendroica petechia aureola.

Pair of Nazca boobies (Sula granti) in a field of flowering goat’s head. Genovesa Island, Galápagos.

Nazca boobies (Sula granti) courting in a field of goat’s head. Genovesa Island, Galápagos.

Nazca booby (Sula granti) with STOP sign. Isla Genovesa, Galápagos.

Nazca booby (Sula granti). Genovesa Island, Galápagos.

Juvenile red-footed booby (Sula sula). Tower Island, Galápagos, Ecuador.

Red-footed booby near Opuntia. Genovesa, Galápagos Islands.

Red-footed booby (Sula sula) on a branch. Isla Genovesa, Galápagos.

Jamaican feverplant (Tribulus cistoides). Isla Bartolomé, Galápagos.

Isla Genovesa, Galápagos.

Flowering goat’s head (Tribulus terrestris). Genovesa, Galápagos Islands, Ecuador.

Isla Isabela, Galapagos., Galápagos.

Galápagos carpenter bee (Xylocopa darwini) on goat’s head (Tribulus terrestris). Newly introduced to Genovesa Island. Galápagos Islands, Ecuador.

Galápagos sea lions (Zalophus wollebaeki) napping on the beach. Tower Island, Galápagos, Ecuador.

Galapagos sea lion (Zalophus wollebaeki) pup nursing. Isla Santiago, Galapagos.

Young Galápagos sea lion (Zalophus wollebaeki) on red sand beach. Isla Rábida, Galápagos.

Galápagos sea lion (Zalophus wollebaeki). Isla Rabida, Galápagos.

Galápagos sea lion (Zalophus wollebaeki) blowing bubbles. Isla Fernandina, Galápagos.

Galápagos dove (Zenaida galapagoensis). Genovesa, Galápagos.